Professional Title
Professor of Biology and the Large Lakes Observatory, Professor Emeritus
Ph.D. in Zoology, 1971, Duke UniversityÂ
Nutrient and carbon cycling in aquatic systems; limnology and paleolimnology of the African Great Lakes; phytoplankton-nutrient interactions; nutrient biogeochemistry; stable isotope applications to ecology; Hg and organochlorine contamination of food webs.